Most of the animal depictions that have made their way ... Dr Grollemond says the purpose of marginalia has been long debated ...

A study of 16 Medieval manuscripts in Clairvaux Abbey in France found they were bound in a surprising material: sealskin.

“The seal was rarely depicted in medieval art and did not hold a place in heraldry, which developed at that time. This lack of representation might have contributed to a detachment from the animal’s ...